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Cyrillaceae family

Swamp Titi Cyrilla racemiflora

Native
Deer favourite — Rated highly palatable to browsing deer — so where deer are around, expect this one to get nibbled. It's a documented preference, not a certainty: pressure swings with the place and season, and a fenced or low-deer yard may be untouched.
Also known as: Leatherwood · Swamp Cyrilla · Titi

Swamp Titi is a perennial tree native to the lower 48 states and Puerto Rico. It grows to 30 ft and blooms Jul in full sun, with brown fruit.

More about this plant

Cyrilla racemiflora, the sole species in the genus Cyrilla, is a flowering plant in the family Cyrillaceae, native to warm temperate to tropical regions of the Americas, from the southeastern United States, south through the Caribbean, Mexico and Central America to northern Brazil and Venezuela in South America. Common names include swamp cyrilla, swamp titi, palo colorado, red titi, black titi, white titi, leatherwood, ironwood, he-huckleberry, and myrtle. Wikipedia →

Growing & care

USDA PLANTS · TRY
Conditions
Sources · Conditions
USDA PLANTS — Sun
USDA — SoilPH — Soil pH
USDA PLANTS (DwCA, Zenodo 17903503) — Fertility Requirement / Soil Adaptation — Fertility need · Adapts to
USDA — Temp-Min °F — Hardiness
Sun
Full sun
Soil pH
4.9–6.8
Fertility need
High
Adapts to
Coarse (sandy), Medium (loam), Fine (clay)
Hardiness
USDA zone 7+
Size & form
Sources · Size & form
USDA PLANTS (DwCA, Zenodo 17903503) — Height, Mature — Height
USDA PLANTS (DwCA, Zenodo 17903503) — Planting Density (per acre) — Spacing
USDA PLANTS (DwCA, Zenodo 17903503) — Growth Rate / Spread / Growth Form / Lifespan / Active Growth Period / Seed Period — Spread · Growth rate · Growth form · Active growth
USDA PLANTS — Lifespan
TRY Plant Trait Database (CC BY 4.0) · USDA PLANTS — Foliage
USDA PLANTS (DwCA, Zenodo 17903503) — Fruit/Seed Color / Fruit Persistence — Fruit
McKenzie et al. 2025 — community-science image analysis (MIT) — Flower colour
Lifespan: USDA’s lifespan scale for trees: short = under ~100 years · moderate = 100–250 · long = over 250. Balsam fir and other short-lived conifers are genuinely short for a tree, not short like an annual.
Height
30 ft
Spacing
6–12 ft apart from USDA planting density
Spread
Moderate
Growth rate
Moderate
Growth form
Thicket-forming
Lifespan
Perennial · short-lived
Foliage
Evergreen broadleaf · coarse texture
Active growth
Spring & summer
Fruit
Brown
Flower colour
White AI AI image-analysis of community-science photos (~87% expert agreement) — not a botanical record
In the garden
Sources · In the garden
USDA PLANTS (DwCA, Zenodo 17903503) — Propagation Method / Commercial Availability — Propagate by · In the trade
USDA PLANTS (DwCA, Zenodo 17903503) — Germination (cold stratification) — Seed starting
USDA PLANTS — Seed Period — Seeds ripen
USDA PLANTS (DwCA, Zenodo 17903503) — Foliage Texture / Browse Palatability / Resprout — Deer browsing · Resprouts if cut
Propagate by
Seed, Cuttings
Seed starting
Needs cold stratification a cold-moist spell before it germinates
Seeds ripen
Summer – Autumn seed-collection / harvest window
In the trade
No known commercial source
Deer browsing
High browsed readily
Resprouts if cut
Yes regrows after top-kill
Sub-canopy / understory layer — Sits in the understory of a layered food forest or polyculture.Open guide →
Rain garden / bioswale — Rated FACW on the wetland list — tolerates wet feet, so it suits the soggy zone of a rain garden or bioswale that catches and filters runoff.Open guide →
derived roles
AI = read by an AI vision model · DERIVED = a computed estimate, not a direct measurement. The “How we know this” section below details each.
The garden year bloom → fruit → fall colour
J
F
M
A
M
J
J
A
S
O
N
D
Bloom
Bloom (the flower's colour)
Bloom · Jul — USDA PLANTS (DwCA, Zenodo 17903503) — BloomPeriod
When to sow · for your area

Sow timing keys off your local last- and first-frost dates.

Species characteristics from USDA PLANTS (public domain) + TRY (CC BY) — general guidance, not a guarantee for your exact site. Deer "browsing" is documented palatability, not a deer-proof claim.

Wildlife & pollinators

How pollinator value is scored →
❧ Caterpillar hosts Documented caterpillar host

Recorded feeding on Cyrilla in North America, including:

Incisalia henrici · gossamer-winged butterfly ✦
Named species (a documented Nearctic sample, not exhaustive) from NHM HOSTS (CC0). ✦ = recorded on this exact species.
✦ Bees 14 bee visitors
Visitor records (observations, not exhaustive) from Noori et al. 2026 (CC BY 4.0) · GloBI → · ✦ = pollen-specialist bee (Smith et al. 2024)
Wildlife & visitors 5 nectaring

Open records of who else uses Swamp Titi — a generalist food-web signal, kept separate from the keystone Ecological Value.

Interaction records (observations, not exhaustive) from GloBI → (CC0). Counts are distinct species; names are the most-recorded. Common names from Wikidata (CC0).
Species thumbnails re-hosted from iNaturalist — Creative Commons, credited per image (hover for credit). Click any species to see it on iNaturalist. Not exhaustive; many taxa have no openly-licensed photo yet.
Frequently grows with

Across 390 U.S. vegetation-survey plots that recorded Swamp Titi, these catalog species turned up most often in the same plot — a real field co-occurrence signal, not a planting prescription.

Blackgum · 72% Red Maple · 68% Laurel Greenbrier · 53% Sweetgum · 51% Sweetbay · 48% Swamp Bay · 46% Eastern Poison Ivy · 46% Fetterbush Lyonia · 45%
"%" = share of this plant's plots that also held the species. From sPlotOpen → (Sabatini et al. 2021, CC BY 4.0) — open vegetation plots: wild community composition, not garden design.
How we know this (4) Methods & honest limits
Leaf build Direct fact

We read each species’ measured specific leaf area (leaf area per unit dry weight) from the Global Spectrum dataset and band it: thin & fast (high SLA), medium, or thick & tough (low SLA). The global median is about 10 mm²/mg.

Honest limits: A species-mean from pooled measurements — individual plants vary with light and site. A broad strategy signal, not a precise per-plant figure.

Díaz, S. et al. (2022) The global spectrum of plant form and function: enhanced species-level trait dataset. Scientific Data 9:755.
Photosynthesis Direct fact

A recorded categorical fact: each species is tagged C3 (standard), C4 (heat/water-efficient) or CAM (succulent, night-time CO₂ uptake) — or a facultative combination. We only show a trait card for the noteworthy C4/CAM cases; C3 is the unremarkable majority, kept in the data but not surfaced as a card.

Kattge, J. et al. TRY plant trait database — Categorical Traits Dataset (2012).
Climate niche (heat tolerance & native rainfall) Derived

We take the plant’s U.S. county range and look up each county’s long-term climate, then summarise: native rainfall = the median annual rainfall across its counties; heat tolerance = the warm end (90th percentile) of average temperature across its range (mirroring how our cold-hardiness floor uses an extreme). Counties + medians blunt the roadside/observer bias that makes point-level climate unreliable. Needs at least five placeable counties.

Honest limits: A realized, sampling-biased niche (where it has been recorded, not its physiological optimum), and county climate is coarse — large Western counties span deserts and mountains. Derived guide, never a measured fact.

Realized-niche / climate-envelope approach (Pearson & Dawson 2003; Soberón 2007). Climate: NOAA NCEI nClimDiv county normals (1991–2020).
Flower colour Derived

McKenzie et al. assigned each species a flower colour with a GPT-4V vision model over iNaturalist photos. We use the confident tier plus a separately-labelled lower-confidence top-up (mostly inconspicuous green/brown flowers), and render it as a small tint — never as an asserted fact, and kept out of the written synopsis.

Honest limits: AI image inference (~87% expert agreement on the confident tier); a decorative, confidence-tiered indicator, not a measured trait.

McKenzie, P., Berardi, A.E., Hopkins, R. (2025). flower_color_phenology (MIT).
Derived values are computed from open data by a published method — labelled, cited, and never shown as a direct observation. Full methodology →
Sources for this entry (37) Open & cited
[01] Scientific name & family — USDA PLANTS (DwCA, Zenodo 17903503)
[02] Growth habit & duration — USDA PLANTS (DwCA, Zenodo 17903503)
[03] Native status & distribution — USDA PLANTS (DwCA, Zenodo 17903503)
[04] Common name — USDA PLANTS (via GBIF)
[17] Invasive / introduced status — USDA PLANTS (DwCA, Zenodo 17903503) — native status
[18] Wetland indicator — USACE National Wetland Plant List (2022)
[20] Description — Wikipedia (CC BY-SA)
[21] Ecological value — GloBI
[22] Conservation rank — NatureServe Explorer (CC BY)
[24] Functional traits — TRY Plant Trait Database (CC BY 4.0)
[25] Photosynthetic pathway — TRY Plant Trait Database (CC BY 4.0)
[26] Climate niche — Derived — realized climate niche from U.S. county occurrences (GBIF) × NOAA NCEI county climate normals 1991–2020 (public domain)
[27] Wildlife & visitors — GloBI — Global Biotic Interactions (CC0)
[31] Other common names — Wikidata (CC0)
[33] Bee specialists — Smith et al. 2024 (Zenodo, CC BY 4.0)
[34] Flower-visitor value (derived) — Noori et al. 2026 (CC BY 4.0) · GloBI
[35] Caterpillar species — NHM HOSTS (CC0)
[36] County range — GBIF Open Data — open-licensed occurrences (CC0/CC-BY) → county
Spot an error or have a better photo? Every field links to its source, and the whole dataset is open.
Cite this page Open data, please attribute

PlantKey’s data is open under CC BY-SA 4.0 — free to reuse and adapt, with attribution and the same licence. Photos keep their own per-image licence + credit (see Sources above).

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